AMPHIBIAWEB

Dicroglossidae

207 species in 14 genera

Commonly Called Forked-Tongue Frogs, Cricket Frogs


Fejervarya limnocharis
Photo by Martin Pickersgill
(Click for family gallery)

This large family has a diversity of forms and natural history modes; likewise it has had a sometimes confusing but active taxonomic history. Its genera are split into two subfamilies, Dicroglossinae (with most of the species) and Occidozyginae. Previously, these species were included in Ranidae and a majority of species were assigned to the genus Fejervarya. In 2018, Fejervarya was proposed to be limited to the Southeast Asia species with the resurrected genus Minervarya restricted to South Asia (Sanchez et al. 2018). Most authors agree that more species are yet to be described in this family.

They range from sub-Saharan Africa south to South Africa east through western Asia, South Asia (India), China, southeast Asia, including Malay peninsula, Indonesia, New Guinea, the Philippines and Japan.

Written by AmphibiaWeb

Notable Family Characteristics

  • Terrestrial and aquatic species, found in a diversity of habitats
  • Both egg-laying and direct developing species
  • Diverse reproductive modes including uncommon inguinal amplexus in Occidozyga sumatrana
  • Includes crab-eating frog Fejervarya cancrivora with unique specializations for living in brackish mangroves
  • Includes burrowing species like Sphaerotheca breviceps, which appear above ground after heavy rains
  • Distributed through sub-Saharan Africa, western Asia, South Asia (India), Southeast Asia, Philippines, New Guinea and Japan

Relevant Reference

Kurabayashi, A., M. Kuramoto, S. H. Joshy, and M. Sumida. 2005. Molecular phylogeny of the ranid frogs from Southwest India based on the mitochondrial ribosomal RNA gene sequences. Zoological Science. Tokyo 22: 525–534.

Roelants, K., J.-p. Jiang, and F. Bossuyt. 2004. Endemic ranid (Amphibia: Anura) genera in southern mountain ranges of the Indian subcontinent represent ancient frog lineages: evidence from the molecular data. Molecular Phylogenetics and Evolution 31: 730–740.

Sanchez, E., S. D. Biju, M. M. Islam, M. K. Hasan, A. Ohler, M. Vences, and A. Kurabayashi. 2018. Phylogeny and classification of fejervaryan frogs (Anura: Dicroglossidae). Salamandra 54: 109–116.

Ohler, A., and A. Dubois. 2006. Phylogenetic relationships and generic taxonomy of the tribe Paini (Amphibia, Anura, Ranidae, Dicroglossinae) with diagnoses of two new genera. Zoosystema. Paris 28: 769–784.

Subfamily Dicroglossinae (192 species)

Genus Allopaa (2 species)    [subfamily Dicroglossinae]
Allopaa barmoachensis no AmphibiaWeb account no photos no sound/video
Allopaa hazarensis AmphibiaWeb account photos no sound/video

Genus Chrysopaa (1 species)    [subfamily Dicroglossinae]
Chrysopaa sternosignata AmphibiaWeb account photos no sound/video

Genus Euphlyctis (8 species)    [subfamily Dicroglossinae]
Euphlyctis aloysii no AmphibiaWeb account photos no sound/video
Euphlyctis cyanophlyctis AmphibiaWeb account photos sound/video
Euphlyctis ehrenbergii no AmphibiaWeb account photos no sound/video
Euphlyctis ghoshi no AmphibiaWeb account no photos no sound/video
Euphlyctis hexadactylus AmphibiaWeb account photos no sound/video
Euphlyctis kalasgramensis no AmphibiaWeb account no photos no sound/video
Euphlyctis karaavali no AmphibiaWeb account no photos no sound/video
Euphlyctis mudigere no AmphibiaWeb account no photos no sound/video

Genus Fejervarya (17 species)    [subfamily Dicroglossinae]
Fejervarya altilabris no AmphibiaWeb account no photos no sound/video
Fejervarya assimilis no AmphibiaWeb account no photos no sound/video
Fejervarya brama no AmphibiaWeb account no photos no sound/video
Fejervarya cancrivora AmphibiaWeb account photos sound/video
Fejervarya frithi no AmphibiaWeb account no photos no sound/video
Fejervarya iskandari no AmphibiaWeb account no photos no sound/video
Fejervarya kawamurai no AmphibiaWeb account no photos no sound/video
Fejervarya limnocharis AmphibiaWeb account photos sound/video
Fejervarya moodiei no AmphibiaWeb account photos no sound/video
Fejervarya multistriata no AmphibiaWeb account no photos no sound/video
Fejervarya orissaensis no AmphibiaWeb account no photos no sound/video
Fejervarya pulla no AmphibiaWeb account no photos no sound/video
Fejervarya raja no AmphibiaWeb account no photos no sound/video
Fejervarya sakishimensis no AmphibiaWeb account no photos no sound/video
Fejervarya triora no AmphibiaWeb account no photos no sound/video
Fejervarya verruculosa no AmphibiaWeb account no photos no sound/video
Fejervarya vittigera no AmphibiaWeb account no photos no sound/video

Genus Hoplobatrachus (5 species)    [subfamily Dicroglossinae]
Hoplobatrachus crassus AmphibiaWeb account photos no sound/video
Hoplobatrachus litoralis no AmphibiaWeb account photos no sound/video
Hoplobatrachus occipitalis AmphibiaWeb account photos sound/video
Hoplobatrachus rugulosus no AmphibiaWeb account photos no sound/video
Hoplobatrachus tigerinus AmphibiaWeb account photos no sound/video

Genus Limnonectes (70 species)    [subfamily Dicroglossinae]
Limnonectes acanthi no AmphibiaWeb account no photos no sound/video
Limnonectes arathooni no AmphibiaWeb account no photos no sound/video
Limnonectes asperatus no AmphibiaWeb account no photos no sound/video
Limnonectes bannaensis no AmphibiaWeb account no photos no sound/video
Limnonectes blythii no AmphibiaWeb account photos no sound/video
Limnonectes cintalubang AmphibiaWeb account no photos no sound/video
Limnonectes coffeatus no AmphibiaWeb account no photos no sound/video
Limnonectes dabanus no AmphibiaWeb account photos no sound/video
Limnonectes dammermani no AmphibiaWeb account photos no sound/video
Limnonectes diuatus no AmphibiaWeb account no photos no sound/video
Limnonectes doriae no AmphibiaWeb account no photos no sound/video
Limnonectes ferneri AmphibiaWeb account no photos no sound/video
Limnonectes finchi no AmphibiaWeb account photos no sound/video
Limnonectes fragilis no AmphibiaWeb account photos no sound/video
Limnonectes fujianensis no AmphibiaWeb account no photos no sound/video
Limnonectes grunniens no AmphibiaWeb account no photos no sound/video
Limnonectes gyldenstolpei no AmphibiaWeb account photos no sound/video
Limnonectes hascheanus no AmphibiaWeb account photos sound/video
Limnonectes heinrichi no AmphibiaWeb account no photos no sound/video
Limnonectes hikidai no AmphibiaWeb account no photos no sound/video
Limnonectes ibanorum no AmphibiaWeb account photos no sound/video
Limnonectes ingeri no AmphibiaWeb account photos no sound/video
Limnonectes isanensis no AmphibiaWeb account no photos no sound/video
Limnonectes jarujini no AmphibiaWeb account no photos no sound/video
Limnonectes kadarsani AmphibiaWeb account photos no sound/video
Limnonectes kenepaiensis no AmphibiaWeb account photos no sound/video
Limnonectes khammonensis no AmphibiaWeb account no photos no sound/video
Limnonectes khasianus no AmphibiaWeb account no photos no sound/video
Limnonectes kohchangae no AmphibiaWeb account photos no sound/video
Limnonectes kong no AmphibiaWeb account photos no sound/video
Limnonectes kuhlii no AmphibiaWeb account photos no sound/video
Limnonectes larvaepartus AmphibiaWeb account photos no sound/video
Limnonectes laticeps no AmphibiaWeb account photos no sound/video
Limnonectes lauhachindai no AmphibiaWeb account no photos no sound/video
Limnonectes leporinus no AmphibiaWeb account photos no sound/video
Limnonectes leytensis no AmphibiaWeb account photos no sound/video
Limnonectes limborgi no AmphibiaWeb account no photos no sound/video
Limnonectes liui no AmphibiaWeb account photos sound/video
Limnonectes longchuanensis no AmphibiaWeb account no photos no sound/video
Limnonectes macrocephalus no AmphibiaWeb account no photos no sound/video
Limnonectes macrodon no AmphibiaWeb account photos no sound/video
Limnonectes macrognathus no AmphibiaWeb account no photos no sound/video
Limnonectes magnus no AmphibiaWeb account no photos no sound/video
Limnonectes malesianus no AmphibiaWeb account photos no sound/video
Limnonectes mawlyndipi no AmphibiaWeb account no photos no sound/video
Limnonectes megastomias AmphibiaWeb account no photos no sound/video
Limnonectes micrixalus no AmphibiaWeb account no photos no sound/video
Limnonectes microdiscus no AmphibiaWeb account no photos no sound/video
Limnonectes microtympanum no AmphibiaWeb account no photos no sound/video
Limnonectes modestus no AmphibiaWeb account no photos no sound/video
Limnonectes namiyei AmphibiaWeb account no photos no sound/video
Limnonectes nguyenorum no AmphibiaWeb account no photos no sound/video
Limnonectes nitidus no AmphibiaWeb account no photos no sound/video
Limnonectes palavanensis no AmphibiaWeb account photos no sound/video
Limnonectes paramacrodon no AmphibiaWeb account photos no sound/video
Limnonectes parvus no AmphibiaWeb account no photos no sound/video
Limnonectes plicatellus no AmphibiaWeb account no photos no sound/video
Limnonectes poilani no AmphibiaWeb account photos no sound/video
Limnonectes quangninhensis no AmphibiaWeb account no photos no sound/video
Limnonectes rhacodus no AmphibiaWeb account no photos no sound/video
Limnonectes selatan no AmphibiaWeb account no photos no sound/video
Limnonectes shompenorum no AmphibiaWeb account no photos no sound/video
Limnonectes sinuatodorsalis AmphibiaWeb account no photos no sound/video
Limnonectes sisikdagu no AmphibiaWeb account no photos no sound/video
Limnonectes taylori no AmphibiaWeb account no photos no sound/video
Limnonectes timorensis no AmphibiaWeb account no photos no sound/video
Limnonectes tweediei no AmphibiaWeb account no photos no sound/video
Limnonectes utara no AmphibiaWeb account no photos no sound/video
Limnonectes visayanus no AmphibiaWeb account photos no sound/video
Limnonectes woodworthi no AmphibiaWeb account no photos no sound/video

Genus Minervarya (36 species)    [subfamily Dicroglossinae]
Minervarya andamanensis no AmphibiaWeb account photos no sound/video
Minervarya asmati no AmphibiaWeb account no photos no sound/video
Minervarya brevipalmata no AmphibiaWeb account no photos no sound/video
Minervarya caperata no AmphibiaWeb account photos no sound/video
Minervarya cepfi no AmphibiaWeb account photos no sound/video
Minervarya chiangmaiensis no AmphibiaWeb account no photos no sound/video
Minervarya chilapata AmphibiaWeb account no photos no sound/video
Minervarya dhaka no AmphibiaWeb account no photos no sound/video
Minervarya goemchi no AmphibiaWeb account no photos no sound/video
Minervarya gomantaki no AmphibiaWeb account photos no sound/video
Minervarya granosa no AmphibiaWeb account photos no sound/video
Minervarya greenii AmphibiaWeb account photos no sound/video
Minervarya jhilmilensis no AmphibiaWeb account no photos no sound/video
Minervarya kadar no AmphibiaWeb account no photos no sound/video
Minervarya kalinga no AmphibiaWeb account no photos no sound/video
Minervarya keralensis no AmphibiaWeb account no photos no sound/video
Minervarya kirtisinghei AmphibiaWeb account photos no sound/video
Minervarya krishnan no AmphibiaWeb account no photos no sound/video
Minervarya kudremukhensis no AmphibiaWeb account photos no sound/video
Minervarya manoharani no AmphibiaWeb account no photos no sound/video
Minervarya modesta no AmphibiaWeb account no photos no sound/video
Minervarya mudduraja no AmphibiaWeb account photos no sound/video
Minervarya murthii no AmphibiaWeb account no photos no sound/video
Minervarya mysorensis no AmphibiaWeb account no photos no sound/video
Minervarya neilcoxi no AmphibiaWeb account no photos no sound/video
Minervarya nepalensis no AmphibiaWeb account no photos no sound/video
Minervarya nicobariensis no AmphibiaWeb account no photos no sound/video
Minervarya nilagirica no AmphibiaWeb account photos no sound/video
Minervarya parambikulamana no AmphibiaWeb account no photos no sound/video
Minervarya pierrei no AmphibiaWeb account photos no sound/video
Minervarya rufescens no AmphibiaWeb account photos no sound/video
Minervarya sahyadris no AmphibiaWeb account photos no sound/video
Minervarya sauriceps no AmphibiaWeb account no photos no sound/video
Minervarya sengupti no AmphibiaWeb account no photos no sound/video
Minervarya syhadrensis AmphibiaWeb account photos no sound/video
Minervarya teraiensis no AmphibiaWeb account photos no sound/video

Genus Nannophrys (4 species)    [subfamily Dicroglossinae]
Nannophrys ceylonensis AmphibiaWeb account photos sound/video
Nannophrys guentheri AmphibiaWeb account no photos no sound/video
Nannophrys marmorata AmphibiaWeb account photos no sound/video
Nannophrys naeyakai AmphibiaWeb account photos no sound/video

Genus Nanorana (29 species)    [subfamily Dicroglossinae]
Nanorana aenea no AmphibiaWeb account photos no sound/video
Nanorana annandalii no AmphibiaWeb account no photos no sound/video
Nanorana arnoldi no AmphibiaWeb account no photos no sound/video
Nanorana blanfordii no AmphibiaWeb account no photos no sound/video
Nanorana chayuensis no AmphibiaWeb account photos no sound/video
Nanorana conaensis no AmphibiaWeb account photos no sound/video
Nanorana delacouri no AmphibiaWeb account no photos no sound/video
Nanorana ercepeae no AmphibiaWeb account no photos no sound/video
Nanorana feae no AmphibiaWeb account no photos no sound/video
Nanorana gammii no AmphibiaWeb account no photos no sound/video
Nanorana kangxianensis no AmphibiaWeb account no photos no sound/video
Nanorana liebigii no AmphibiaWeb account no photos no sound/video
Nanorana maculosa no AmphibiaWeb account no photos no sound/video
Nanorana medogensis no AmphibiaWeb account photos no sound/video
Nanorana minica no AmphibiaWeb account photos no sound/video
Nanorana mokokchungensis no AmphibiaWeb account no photos no sound/video
Nanorana parkeri no AmphibiaWeb account photos no sound/video
Nanorana phrynoides no AmphibiaWeb account no photos no sound/video
Nanorana pleskei no AmphibiaWeb account photos no sound/video
Nanorana polunini no AmphibiaWeb account no photos no sound/video
Nanorana quadranus no AmphibiaWeb account no photos no sound/video
Nanorana rarica no AmphibiaWeb account no photos no sound/video
Nanorana rostandi no AmphibiaWeb account no photos no sound/video
Nanorana sichuanensis no AmphibiaWeb account no photos no sound/video
Nanorana taihangnica no AmphibiaWeb account no photos no sound/video
Nanorana unculuanus no AmphibiaWeb account no photos no sound/video
Nanorana ventripunctata no AmphibiaWeb account photos no sound/video
Nanorana vicina AmphibiaWeb account photos no sound/video
Nanorana yunnanensis AmphibiaWeb account no photos no sound/video

Genus Ombrana (1 species)    [subfamily Dicroglossinae]
Ombrana sikimensis no AmphibiaWeb account no photos no sound/video

Genus Quasipaa (10 species)    [subfamily Dicroglossinae]
Quasipaa acanthophora no AmphibiaWeb account no photos no sound/video
Quasipaa boulengeri no AmphibiaWeb account photos no sound/video
Quasipaa courtoisi no AmphibiaWeb account no photos no sound/video
Quasipaa exilispinosa no AmphibiaWeb account photos no sound/video
Quasipaa fasciculispina no AmphibiaWeb account no photos no sound/video
Quasipaa jiulongensis no AmphibiaWeb account no photos no sound/video
Quasipaa shini no AmphibiaWeb account no photos no sound/video
Quasipaa spinosa no AmphibiaWeb account no photos no sound/video
Quasipaa verrucospinosa no AmphibiaWeb account photos no sound/video
Quasipaa yei no AmphibiaWeb account no photos no sound/video

Genus Sphaerotheca (9 species)    [subfamily Dicroglossinae]
Sphaerotheca breviceps AmphibiaWeb account photos no sound/video
Sphaerotheca dobsonii no AmphibiaWeb account photos no sound/video
Sphaerotheca leucorhynchus no AmphibiaWeb account no photos no sound/video
Sphaerotheca maskeyi no AmphibiaWeb account no photos no sound/video
Sphaerotheca pashchima no AmphibiaWeb account photos no sound/video
Sphaerotheca pluvialis no AmphibiaWeb account no photos no sound/video
Sphaerotheca rolandae AmphibiaWeb account photos no sound/video
Sphaerotheca strachani no AmphibiaWeb account no photos no sound/video
Sphaerotheca swani no AmphibiaWeb account no photos no sound/video
Subfamily Occidozyginae (15 species)

Genus Ingerana (3 species)    [subfamily Occidozyginae]
Ingerana borealis no AmphibiaWeb account photos no sound/video
Ingerana charlesdarwini no AmphibiaWeb account no photos no sound/video
Ingerana tenasserimensis no AmphibiaWeb account no photos no sound/video

Genus Occidozyga (12 species)    [subfamily Occidozyginae]
Occidozyga baluensis no AmphibiaWeb account photos no sound/video
Occidozyga celebensis no AmphibiaWeb account photos no sound/video
Occidozyga diminutiva no AmphibiaWeb account no photos no sound/video
Occidozyga floresiana no AmphibiaWeb account no photos no sound/video
Occidozyga laevis AmphibiaWeb account photos sound/video
Occidozyga lima no AmphibiaWeb account photos no sound/video
Occidozyga magnapustulosa no AmphibiaWeb account no photos no sound/video
Occidozyga martensii no AmphibiaWeb account photos sound/video
Occidozyga semipalmata no AmphibiaWeb account photos no sound/video
Occidozyga sumatrana no AmphibiaWeb account photos no sound/video
Occidozyga tompotika no AmphibiaWeb account no photos no sound/video
Occidozyga vittata no AmphibiaWeb account no photos no sound/video


Citation: AmphibiaWeb: Information on amphibian biology and conservation. [web application]. Berkeley, California: AmphibiaWeb. Available: https://amphibiaweb.org/. (Accessed:

AmphibiaWeb's policy on data use.