AmphibiaWeb - Dicroglossidae
AMPHIBIAWEB

Dicroglossidae

223 species in 15 genera

Commonly Called Forked-Tongue Frogs, Cricket Frogs


Fejervarya limnocharis
Photo by Martin Pickersgill
(Click for family gallery)

This large family has a diversity of forms and natural history modes; likewise it has had a sometimes confusing but active taxonomic history. Its genera are split into two subfamilies, Dicroglossinae (with most of the species) and Occidozyginae. Previously, these species were included in Ranidae and a majority of species were assigned to the genus Fejervarya. In 2018, Fejervarya was proposed to be limited to the Southeast Asia species with the resurrected genus Minervarya restricted to South Asia (Sanchez et al. 2018). Most authors agree that more species are yet to be described in this family.

They range from sub-Saharan Africa south to South Africa east through western Asia, South Asia (India), China, southeast Asia, including Malay peninsula, Indonesia, New Guinea, the Philippines and Japan.

Written by AmphibiaWeb

Notable Family Characteristics

  • Terrestrial and aquatic species, found in a diversity of habitats
  • Both egg-laying and direct developing species
  • Diverse reproductive modes including uncommon inguinal amplexus in Occidozyga sumatrana
  • Includes crab-eating frog Fejervarya cancrivora with unique specializations for living in brackish mangroves
  • Includes burrowing species like Sphaerotheca breviceps, which appear above ground after heavy rains
  • Distributed through sub-Saharan Africa, western Asia, South Asia (India), Southeast Asia, Philippines, New Guinea and Japan
Dicroglossidae Richness map

Cartography Credit: Zoe Yoo, UC Berkeley
Range maps sources: AmphibiaWeb, UC Berkeley, and IUCN RedList

Relevant Reference

Kurabayashi, A., M. Kuramoto, S. H. Joshy, and M. Sumida. 2005. Molecular phylogeny of the ranid frogs from Southwest India based on the mitochondrial ribosomal RNA gene sequences. Zoological Science. Tokyo 22: 525–534.

Roelants, K., J.-p. Jiang, and F. Bossuyt. 2004. Endemic ranid (Amphibia: Anura) genera in southern mountain ranges of the Indian subcontinent represent ancient frog lineages: evidence from the molecular data. Molecular Phylogenetics and Evolution 31: 730–740.

Sanchez, E., S. D. Biju, M. M. Islam, M. K. Hasan, A. Ohler, M. Vences, and A. Kurabayashi. 2018. Phylogeny and classification of fejervaryan frogs (Anura: Dicroglossidae). Salamandra 54: 109–116.

Ohler, A., and A. Dubois. 2006. Phylogenetic relationships and generic taxonomy of the tribe Paini (Amphibia, Anura, Ranidae, Dicroglossinae) with diagnoses of two new genera. Zoosystema. Paris 28: 769–784.

Subfamily Dicroglossinae (207 species)

Genus Allopaa (2 species)    [subfamily Dicroglossinae]
Allopaa barmoachensis no account no photos no sound/video
Allopaa hazarensis account photos no sound/video

Genus Chrysopaa (1 species)    [subfamily Dicroglossinae]
Chrysopaa sternosignata account photos no sound/video

Genus Euphlyctis (9 species)    [subfamily Dicroglossinae]
Euphlyctis aloysii no account photos no sound/video
Euphlyctis cyanophlyctis account photos sound/video
Euphlyctis ehrenbergii no account photos no sound/video
Euphlyctis ghoshi no account no photos no sound/video
Euphlyctis hexadactylus account photos no sound/video
Euphlyctis kalasgramensis no account no photos no sound/video
Euphlyctis karaavali no account no photos no sound/video
Euphlyctis kerala no account no photos no sound/video
Euphlyctis mudigere no account no photos no sound/video

Genus Fejervarya (17 species)    [subfamily Dicroglossinae]
Fejervarya altilabris no account no photos no sound/video
Fejervarya assimilis no account no photos no sound/video
Fejervarya brama no account no photos no sound/video
Fejervarya cancrivora account photos sound/video
Fejervarya frithi no account no photos no sound/video
Fejervarya iskandari no account no photos no sound/video
Fejervarya kawamurai no account no photos no sound/video
Fejervarya kupitzi no account no photos no sound/video
Fejervarya limnocharis account photos sound/video
Fejervarya moodiei no account photos no sound/video
Fejervarya multistriata no account photos no sound/video
Fejervarya orissaensis no account no photos no sound/video
Fejervarya pulla no account no photos no sound/video
Fejervarya sakishimensis no account no photos no sound/video
Fejervarya triora no account no photos no sound/video
Fejervarya verruculosa no account no photos no sound/video
Fejervarya vittigera no account no photos no sound/video

Genus Hoplobatrachus (5 species)    [subfamily Dicroglossinae]
Hoplobatrachus crassus account photos no sound/video
Hoplobatrachus litoralis no account photos no sound/video
Hoplobatrachus occipitalis account photos sound/video
Hoplobatrachus rugulosus no account photos no sound/video
Hoplobatrachus tigerinus account photos no sound/video

Genus Limnonectes (77 species)    [subfamily Dicroglossinae]
Limnonectes acanthi no account no photos no sound/video
Limnonectes arathooni no account no photos no sound/video
Limnonectes asperatus no account no photos no sound/video
Limnonectes bannaensis no account no photos no sound/video
Limnonectes beloncioi no account no photos no sound/video
Limnonectes blythii account photos no sound/video
Limnonectes cintalubang account photos no sound/video
Limnonectes coffeatus no account no photos no sound/video
Limnonectes conspicillatus no account no photos no sound/video
Limnonectes dabanus no account photos no sound/video
Limnonectes dammermani no account photos no sound/video
Limnonectes deinodon no account no photos no sound/video
Limnonectes diuatus no account no photos no sound/video
Limnonectes doriae no account no photos no sound/video
Limnonectes fastigatus no account photos no sound/video
Limnonectes ferneri account no photos no sound/video
Limnonectes finchi no account photos no sound/video
Limnonectes fragilis no account photos no sound/video
Limnonectes fujianensis no account photos no sound/video
Limnonectes grunniens no account no photos no sound/video
Limnonectes gyldenstolpei no account photos no sound/video
Limnonectes hascheanus no account photos sound/video
Limnonectes heinrichi no account no photos no sound/video
Limnonectes hikidai no account photos no sound/video
Limnonectes ibanorum no account photos no sound/video
Limnonectes ingeri no account photos no sound/video
Limnonectes isanensis no account no photos no sound/video
Limnonectes jarujini no account no photos no sound/video
Limnonectes kadarsani account photos no sound/video
Limnonectes kenepaiensis no account photos no sound/video
Limnonectes khammonensis no account no photos no sound/video
Limnonectes khasianus no account no photos no sound/video
Limnonectes kiziriani no account photos no sound/video
Limnonectes kohchangae no account photos no sound/video
Limnonectes kong no account photos no sound/video
Limnonectes kuhlii no account photos no sound/video
Limnonectes larvaepartus account photos no sound/video
Limnonectes lauhachindai no account no photos no sound/video
Limnonectes leporinus no account photos no sound/video
Limnonectes leytensis no account photos no sound/video
Limnonectes limborgi no account no photos no sound/video
Limnonectes liui no account photos sound/video
Limnonectes longchuanensis no account no photos no sound/video
Limnonectes macrocephalus no account no photos no sound/video
Limnonectes macrodon no account photos no sound/video
Limnonectes macrognathus no account no photos no sound/video
Limnonectes magnus no account no photos no sound/video
Limnonectes malesianus no account photos no sound/video
Limnonectes mawlyndipi no account no photos no sound/video
Limnonectes megastomias account no photos no sound/video
Limnonectes micrixalus no account no photos no sound/video
Limnonectes microdiscus no account no photos no sound/video
Limnonectes microtympanum no account no photos no sound/video
Limnonectes modestus no account no photos no sound/video
Limnonectes namiyei account no photos no sound/video
Limnonectes nguyenorum no account no photos no sound/video
Limnonectes nitidus no account photos no sound/video
Limnonectes palavanensis account photos no sound/video
Limnonectes paramacrodon no account photos no sound/video
Limnonectes parvus no account no photos no sound/video
Limnonectes phuyenensis no account photos no sound/video
Limnonectes plicatellus no account photos no sound/video
Limnonectes poilani no account photos no sound/video
Limnonectes pseudodoriae no account no photos no sound/video
Limnonectes quangninhensis no account no photos no sound/video
Limnonectes rhacodus no account no photos no sound/video
Limnonectes savan no account no photos no sound/video
Limnonectes selatan no account photos no sound/video
Limnonectes shompenorum no account no photos no sound/video
Limnonectes sinuatodorsalis account no photos no sound/video
Limnonectes sisikdagu no account no photos no sound/video
Limnonectes taylori no account no photos no sound/video
Limnonectes timorensis no account photos no sound/video
Limnonectes tweediei no account no photos no sound/video
Limnonectes utara no account no photos no sound/video
Limnonectes visayanus no account photos no sound/video
Limnonectes woodworthi no account no photos no sound/video

Genus Minervarya (37 species)    [subfamily Dicroglossinae]
Minervarya agricola no account photos no sound/video
Minervarya andamanensis no account photos no sound/video
Minervarya asmati no account no photos no sound/video
Minervarya brevipalmata no account no photos no sound/video
Minervarya cepfi no account photos no sound/video
Minervarya chiangmaiensis no account no photos no sound/video
Minervarya chilapata account no photos no sound/video
Minervarya dhaka no account no photos no sound/video
Minervarya goemchi no account no photos no sound/video
Minervarya gomantaki no account photos no sound/video
Minervarya greenii account photos no sound/video
Minervarya jhilmilensis no account no photos no sound/video
Minervarya kadar no account no photos no sound/video
Minervarya kalinga no account no photos no sound/video
Minervarya keralensis no account photos no sound/video
Minervarya kirtisinghei account photos no sound/video
Minervarya krishnan no account no photos no sound/video
Minervarya kudremukhensis no account photos no sound/video
Minervarya manoharani no account no photos no sound/video
Minervarya marathi no account no photos no sound/video
Minervarya modesta no account no photos no sound/video
Minervarya muangkanensis no account no photos no sound/video
Minervarya mudduraja no account photos no sound/video
Minervarya murthii no account no photos no sound/video
Minervarya mysorensis no account no photos no sound/video
Minervarya neilcoxi no account no photos no sound/video
Minervarya nepalensis no account photos no sound/video
Minervarya nicobariensis no account no photos no sound/video
Minervarya nilagirica no account photos no sound/video
Minervarya parambikulamana no account no photos no sound/video
Minervarya pierrei no account photos no sound/video
Minervarya rufescens no account photos no sound/video
Minervarya sahyadris no account photos no sound/video
Minervarya sauriceps no account no photos no sound/video
Minervarya sengupti no account no photos no sound/video
Minervarya syhadrensis account photos no sound/video
Minervarya teraiensis no account photos no sound/video

Genus Nannophrys (4 species)    [subfamily Dicroglossinae]
Nannophrys ceylonensis account photos sound/video
Nannophrys guentheri account no photos no sound/video
Nannophrys marmorata account photos no sound/video
Nannophrys naeyakai account photos no sound/video

Genus Nanorana (31 species)    [subfamily Dicroglossinae]
Nanorana aenea no account photos no sound/video
Nanorana annandalii no account no photos no sound/video
Nanorana arnoldi no account no photos no sound/video
Nanorana arunachalensis no account no photos no sound/video
Nanorana blanfordii no account no photos no sound/video
Nanorana chayuensis no account photos no sound/video
Nanorana conaensis no account photos no sound/video
Nanorana ercepeae no account no photos no sound/video
Nanorana feae no account no photos no sound/video
Nanorana gammii no account no photos no sound/video
Nanorana kangxianensis no account no photos no sound/video
Nanorana liebigii no account no photos no sound/video
Nanorana maculosa no account no photos no sound/video
Nanorana medogensis no account photos no sound/video
Nanorana minica no account photos no sound/video
Nanorana mokokchungensis no account no photos no sound/video
Nanorana parkeri account photos no sound/video
Nanorana phrynoides no account no photos no sound/video
Nanorana pleskei no account photos no sound/video
Nanorana polunini no account no photos no sound/video
Nanorana quadranus no account no photos no sound/video
Nanorana rarica no account no photos no sound/video
Nanorana rostandi no account no photos no sound/video
Nanorana sichuanensis no account no photos no sound/video
Nanorana taihangnica no account no photos no sound/video
Nanorana unculuanus no account no photos no sound/video
Nanorana ventripunctata no account photos no sound/video
Nanorana vicina account photos no sound/video
Nanorana xuelinensis no account no photos no sound/video
Nanorana yunnanensis account photos no sound/video
Nanorana zhaoermii no account no photos no sound/video

Genus Ombrana (1 species)    [subfamily Dicroglossinae]
Ombrana sikimensis no account no photos no sound/video

Genus Quasipaa (12 species)    [subfamily Dicroglossinae]
Quasipaa acanthophora no account no photos no sound/video
Quasipaa boulengeri no account photos no sound/video
Quasipaa courtoisi no account no photos no sound/video
Quasipaa delacouri no account no photos no sound/video
Quasipaa exilispinosa no account photos sound/video
Quasipaa fasciculispina no account no photos no sound/video
Quasipaa jiulongensis no account no photos no sound/video
Quasipaa robertingeri no account no photos no sound/video
Quasipaa shini no account no photos no sound/video
Quasipaa spinosa no account photos no sound/video
Quasipaa verrucospinosa no account photos no sound/video
Quasipaa yei no account no photos no sound/video

Genus Sphaerotheca (11 species)    [subfamily Dicroglossinae]
Sphaerotheca bengaluru no account no photos no sound/video
Sphaerotheca breviceps account photos no sound/video
Sphaerotheca dobsonii no account photos no sound/video
Sphaerotheca leucorhynchus no account no photos no sound/video
Sphaerotheca magadha no account no photos no sound/video
Sphaerotheca maskeyi no account photos no sound/video
Sphaerotheca pashchima no account photos no sound/video
Sphaerotheca pluvialis no account photos no sound/video
Sphaerotheca rolandae account photos no sound/video
Sphaerotheca strachani no account no photos no sound/video
Sphaerotheca swani no account photos no sound/video
Subfamily Occidozyginae (16 species)

Genus Ingerana (3 species)    [subfamily Occidozyginae]
Ingerana borealis no account photos no sound/video
Ingerana charlesdarwini account no photos no sound/video
Ingerana tenasserimensis no account no photos no sound/video

Genus Occidozyga (2 species)    [subfamily Occidozyginae]
Occidozyga berbeza no account no photos no sound/video
Occidozyga lima no account photos no sound/video

Genus Phrynoglossus (11 species)    [subfamily Occidozyginae]
Phrynoglossus baluensis no account photos no sound/video
Phrynoglossus celebensis no account photos no sound/video
Phrynoglossus diminutivus no account no photos no sound/video
Phrynoglossus floresianus no account no photos no sound/video
Phrynoglossus laevis account photos sound/video
Phrynoglossus magnapustulosus no account no photos no sound/video
Phrynoglossus martensii no account photos sound/video
Phrynoglossus myanhessei no account no photos no sound/video
Phrynoglossus semipalmatus no account photos no sound/video
Phrynoglossus sumatranus no account photos no sound/video
Phrynoglossus tompotika no account no photos no sound/video


Citation: AmphibiaWeb: Information on amphibian biology and conservation. [web application]. Berkeley, California: AmphibiaWeb. Available: https://amphibiaweb.org/. (Accessed:

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