AMPHIBIAWEB

Contributors to AmphibiaWeb
These people generously contributed their knowledge, materials, and time to AmphibiaWeb!

Global Partners
Australia Jean-Marc Hero
Cuba Ansel Fong G.
Madagascar Miguel Vences
Pacific Region Clare Morrison
Spain Jaime Bosch, Ernesto Recuero

Contributors
Franco Andreone Species accounts, Photos
Phil Bishop Species accounts
Ronald Botterweg Updating Species Lists
Carlos Davidson Sound files (Pacific Coast, U. S.)
Ansel Fong G. Species Accounts (Cuba)
Jinzhong Fu Species accounts, Photos
Varad B. Giri Species accounts, Photos
Frank Glaw Species accounts, Photos, Sound files
Margaret Gunzburger Species accounts, Photos
Jean-Marc Hero Species accounts, Photos
Peter Janzen Species accounts, Photos, Sound files
M.S. Khan Species accounts
Sergius L. Kuzmin Species accounts, Photos
Gururaja KV Species accounts, Photos
William P. Leonard Species accounts, Photos
Rafael I. Marquez Photos, Sound files
Masafumi Matsui Species accounts
John Measey Species accounts
Diogo B. Provete Species accounts, Photos
Jean Raffaëlli Species accounts
Jeff Rice/Western Soundscape Archive Frog and salamander call sound files (Western U. S.)
Stephen C. Richter Species accounts, Photos
M.O. Roedel Species accounts
Gonçalo Rosa Species accounts, Photos
Jirka Schmidt Literature Review and Species Updates
Max Sparreboom Species accounts, Photos
Arne Schiřtz Species accounts, Photos
Adam P. Summers Species accounts, Photos
Miguel Vences Species accounts, Photos, Sound files
Luis J. Villanueva-Rivera Species accounts, Photos
Marvalee Wake Species accounts, Photos
Yuezhao Wang Species accounts
Feng Xie Species accounts

Photographers
Mark Aartse-Tuyn Photos
John Abramyan Photos
Aldemar A. Acevedo Photos
Dr. Gururaja K.V. Acharya Photos
Andres Acosta Photos
Dr. Beat Akeret Photos
Elmer C. Aldrich Photos
Ronn Altig Photos
Esteban Alzate Photos
Christine Ambrose Photos
Jean-Louis Amiet Photos
Franco Andreone Photos
Dr. Ariadne Angulo Photos
André Pinassi Antunes Photos
Arachnokulture Photos
Matthew J. Aresco Photos
Nicolas A. Arms Photos
Chris Austin Photos
Robson W. Avila Photos
Andrés Avila Aguilar Photos
Sam Bacchini Photos
Paul Bachhausen Photos
Mark Bakkers Photos
Gustavo A. Ballen Photos
Foster Bam Photos
Rakkasan Barnett Photos
César L. Barrio Amoros Photos
Sean Barry Photos
Marcus Bartelds Photos
Richard D. Bartlett Photos
R.A. Barton Photos
James W. Beck Photos
Albert P. Bekker Photos
Michael Benard Photos
Anna Bennett Photos
Seth B. Benson Photos
Michal Berec Photos
Jamie B. Bettaso Photos
Wouter Beukema Photos
Cathy Bevier Photos
G.K. Bhat Photos
David Bickford Photos
Phil Bishop Photos
Dave Blackburn Photos
John Blenis Photos
Horia Bogdan Photos
Renaud Boistel Photos
Jason Bond Photos
Maciej Bonk Photos
Leonardo Boquimpani-Freitas Photos
Andrew Borcher Photos
Camila Both Photos
Phil Bowles Photos
Eduardo Boza Oviedo Photos
Milo Brichta Photos
D. Broadley Photos
Christopher Bronny Photos
Rafe Brown Photos
Chris Brown Photos
Eugene Bruins Photos
Harold C. Bryant Photos
Dr. Michael Buckmaster Photos
Stephen D. Busack Photos
Kim Cabrera Photos
Rodrigo Cajade Photos
Charles L. Camp Photos
Jonathan Campbell Photos
David C. Cannatella Photos
John Carr Photos
Sergio Potsch Carvalho-e-Silva Photos
Jose Cassimiro Photos
Alessandro Catenazzi Photos
Giovanni Alberto Chaves Portilla Photos
German Chavez Photos
Jing Che Photos
Li Cheng Photos
Hugo Claessen Photos
John Clare Photos
Patrick Coin Photos
Grant Connette Photos
Anita E. Cook Photos
Gerald and Buff Corsi Photos
Ricardo Cossio Photos
Andrew J. Crawford Photos
Paul Crump Photos
Nicole Danos Photos
Biju Das Photos
H. Vannoy Davis Photos
Philip de Pous Photos
A. de Saix Photos
Stephen M. Deban Photos
Maximilian Dehling Photos
Jesse Delia Photos
Dennis Demello Photos
Stephanie Desranleau Photos
Ana Laura Correa and Ismael E. di Tada Photos
K.P. Dinesh Photos
John Dittes Photos
Joseph S. Dixon Photos
Maik Dobiey Photos
Dr. Thomas Doerner Photos
Stefano Doglio Photos
J. Dorda Photos
Joseph Dougherty Photos
Robert B. Douglas Photos
Robert C. Drewes Photos
Carl du Toit Photos
William E. Duellman Photos
E.A. Dunayev Photos
G. Dusej Photos
Jeremiah Easter Photos
Mehregan Ebrahimi Photos
Devin Edmonds Photos
Thomas Eimermacher Photos
Tobias Eisenberg Photos
Kathryn R. Elmer Photos
Frank E. (Ed) Ely Photos
Victor Fabio Luna Photos
Lars Fehlandt Photos
Zach Felix Photos
Danté B Fenolio Photos
Xavier Ferrer Photos
Robert Fewster Photos
Pierre Fidenci Photos
David Fischer Photos
William Flaxington Photos
Ansel Fong Photos
Antoine Fouquet Photos
Raphael Fraga Photos
Alejandro Franco Photos
Michael Frede Photos
Jinzhong Fu Photos
Matthew Fujita Photos
W. Chris Funk Photos
Mario Garcia-Paris Photos
Adrian Garda Photos
Richard Gassaway Photos
Herbert Gasser Photos
Dr. Daniel L. Geiger Photos
Yuri Sheludko and Irina Gerasimenko Photos
Justin Gerlach Photos
C. Giacoma Photos
Ariovaldo Giaretta Photos
Jason Gibson Photos
Graeme R. Gillespie Photos
Varad Giri Photos
Frank Glaw Photos
M. Goed Photos
Felipe Gomes Photos
Lloyd Gomez Photos
Ivan Gomez-Mestre Photos
Dr. Stefan Gorzula Photos
Lucas Grandinetti Photos
Gustavo Grandjean Photos
Jan Grathwohl Photos
Brian Gratwicke Photos
Jose Grau Photos
Hannah Graveling Photos
Michael Graziano Photos
David M. Green Photos
Eli Greenbaum Photos
Harry Greene Photos
Tom Greer Photos
Marguerite Gregory Photos
Edgardo Griffith Photos
Ken Griffiths Photos
Joyce Gross Photos
Marius-Ioan Groza Photos
Juan M. Guayasamin Photos
Serkan Gul Photos
Margaret S. Gunzburger Photos
Rubén Guzmán Photos
Vaclav Gvozdik Photos
Alexander Haas Photos
Celio Haddad Photos
J.G. Hall Photos
Roger Hall Photos
Daniel Bennett/Katie Hampson Photos
James Hanken Photos
Robert W. Hansen Photos
James H. Harding Photos
George W. Hartwell Photos
Dirk Heidbuechel Photos
Jim Hendel Photos
Ralf Hendrix Photos
Vladien Henriquez Photos
Carlos Hernández Photos
Omar Hernández-Ordóńez Photos
Jean-Marc Hero Photos
H.-J. Herrmann Photos
Ngoc Hoang Photos
Walter Hoedl Photos
Kristine Hoffmann Photos
Charles D. Holliger Photos
Robert Howard Photos
Lisa Hug Photos
Alan Hyde Photos
P. Ibisch Photos
Dr. Lloyd Glenn Ingles Photos
Lars L. Iversen Photos
George Jackson Photos
Kate Jackson Photos
Arnaud Jamin Photos
Dr. Peter Janzen Photos
John Jensen Photos
Timothy Johnson Photos
William Jones Photos
K.-H. Jungfer Photos
Hinrich Kaiser Photos
Kevin Kasenchak Photos
John Kast Photos
Ronalda Keith Photos
Andreas Kettenburg Photos
Muhammad Sharif Khan Photos
John Kipping Photos
Ross Knowles Photos
J. Koehler Photos
Philippe Kok Photos
Michelle S. Koo Photos
Iris Koski Photos
Kerry Kriger Photos
Hari Krishnan S. Photos
Brian Kubicki Photos
John P. Clare & Kurt Kunze Photos
Dr. Nick V. Kurzenko Photos
L.N. Kuzmin Photos
Sergius Kuzmin Photos
Dr. Axel Kwet Photos
Enrique La Marca Photos
Usha Lachungpa Photos
Trip Lamb Photos
Victor S. Lamoureux Photos
Louis-M. Landry Photos
Richard Lang Photos
Karia Lara Photos
Malcolm Largen Photos
Shobi Lawalata Photos
Guillermo Lopez Leal Photos
Joel Ledford Photos
Julian C. Lee Photos
Twan Leenders Photos
William Leonard Photos
Mark V. Leppin Photos
Dr. Alan E. Leviton Photos
Paul Licht Photos
Wang Lijun Photos
Tammy Lim Photos
Dong Lin Photos
Harrier Lin Photos
Shannon Lindquist Photos
Rodrigo Lingnau Photos
Nathan Litjens Photos
Spartak Litvinchuk Photos
Mario Lo Valvo Photos
Daniel Loebmann Photos
Ramiro Logares Photos
Mario Lutz Photos
John D. Lynch Photos
Gonçalo M. Rosa Photos
Alberto Maceda Veiga Photos
Robert Macey Photos
Raphael M. Macieira Photos
Natan Maciel Photos
Norio Maeda Photos
Luke Mahler Photos
Samizdat Makein Photos
Shawn Mallan Photos
Raul Maneyro Photos
Dave Mangham Photos
Tim Manolis Photos
Ulrich Manthey Photos
Karina Marques Photos
Eric Marquette Photos
Dr. Rafael I. Marquez Photos
Ole Martin Photos
Patrick Martin Photos
Ińigo Martinez-Solano Photos
Brad Maryan Photos
Dr. Elena Marzona Photos
Rafaqat Masroor Photos
Rubens H. Matsushita Photos
W.C. Matthews Photos
Chris Mattison Photos
Eric Maxwell Photos
Anatoliy Maystrenko Photos
Bejat McCracken Photos
Glenn McCrea Photos
Roy McDiarmid Photos
Jimmy McGuire Photos
John Measey Photos
Dr. Madhava Meegaskumbura Photos
Joseph Mendelson Photos
Vincenzo Mercurio Photos
S.V. Mezhzherin Photos
Susan Middleton Photos
Jeff Miller Photos
Brian Miller Photos
Jessica Miller Photos
Alden H. Miller Photos
Eduardo Minte H. Photos
Jeffrey Mitchell Photos
Mauricio Molina Photos
Brad Moon Photos
Dr. Jiri Moravec Photos
Patrick C. Morgan Photos
Clare Morrison Photos
George Moser Photos
Bill Moses Photos
Albert Most Photos
Omid Mozaffari Photos
Gonzalo R. Mucientes Sandoval Photos
Daniel G. Mulcahy Photos
Arturo Muńoz Photos
Jerry Murray Photos
Gary Nafis Photos
Edward J. Narayan Photos
Nathan Nazdrowicz Photos
Selvino Neckel Photos
Nate Nelson Photos
Hien Ngoc Nguyen Photos
Matthew Niemiller Photos
David Nixon Photos
Andreas Noellert Photos
Christiano Nogueira Photos
Ross Nolly Photos
Bradford Norman Photos
T. Van Nunnery Photos
Wolfgang Ochojski Photos
Annemarie Ohler Photos
Vide Ohlin Photos
Eric Olson Photos
Jo-Ann Ordano Photos
Nikolai Orlov Photos
Dr. Robert T. and Margaret Orr Photos
Diego Ortiz Photos
Thomas Ostrowski Photos
Anaé Otto Photos
Kristiina Ovaska Photos
Maciej Pabijan Photos
Theodore Papenfuss Photos
Duncan Parks Photos
Juan Parra Photos
Gabriela Parra Olea Photos
James L. Patton Photos
Pedro L.V. Peloso Photos
J.L. Perret Photos
Weslei Pertel Photos
Bill Peterman Photos
Michael Peters Photos
Silviu Petrovan Photos
Martin Pickersgill Photos
Patrick Pikacha Photos
Lisa Powers Photos
Rebecca Pradha Photos
Serban Proches Photos
Diogo B. Provete Photos
Robert Puschendorf Photos
Felipe E. Rabanal Gatica Photos
Sindhu Ramchandran Photos
Edoardo Razzetti Photos
Steffen Reichle Photos
Mark Reid Photos
Ksas Rémi Photos
Alan Resetar Photos
AHM Ali Reza Photos
Barry Rice Photos
Christine Richards Photos
Stephen C. Richter Photos
Ignacio de la Riva Photos
Mauricio Rivera Correa Photos
Gustavo Rodriguez Gallegos Photos
M.O. Roedel Photos
Antonio Romano Photos
Santiago Ron Photos
Sean Michael Rovito Photos
Bikramadittya Guha Roy Photos
Liam Ryan Photos
Dr. Paddy Ryan Photos
Mario Sacramento Photos
Richard Sage Photos
Andrea B. Salzman Photos
Miroslav Samardzic Photos
Raquel R. Santos Photos
Gary Sargent Photos
Ryan Sawby Photos
Eduardo Federico Schaefer Photos
Vince Scheidt Photos
Rob Schell Photos
Arne Schiotz Photos
Andreas Schlueter Photos
Kristel Schneider Photos
Sean Schoville Photos
Tiffany Schriever Photos
Aaron Schusteff Photos
UC Berkeley Scientific Photo Lab Photos
Magno Segalla Photos
Fall 2002 Amphibian Decline Seminar at Harvard Photos
Jose M. Serrano Photos
Jackson Shedd Photos
Alison M. Sheehey Photos
E.S. Shepard Photos
David Simi Photos
Dr. Joanna M. Smith Photos
Mirco Sole Photos
Angel Solis Photos
Max Sparreboom Photos
Bill Stagnaro Photos
Robert C. Stebbins Photos
Frank Steinmann Photos
Barry Stephenson Photos
Brett Stevenson Photos
Sam Stewart Photos
Margaret Stewart Photos
Dr. Matthias Stoeck Photos
Tracy I. Storer Photos
R. Straatman Photos
Hanna Strauss Photos
Jeet Sukumaran Photos
John Sullivan Photos
Adam P. Summers Photos
Javier Sunyer Photos
Kelly Sweet Photos
Andrea Swei Photos
Tigran Tadevosyan Photos
Mills Tandy Photos
Benjamin Tapley Photos
D.N. Tarkhnishvili Photos
John H. Tashijian Photos
Frank Teigler Photos
Bernardo Franco da Veiga Teixeira Photos
Mauro Teixeira Jr. Photos
David A. Tharp Photos
Chris Thawley Photos
B. Thiesmeier Photos
Stephen G. Tilley Photos
Boris I. Timofeev Photos
Geordie Torr Photos
Russell Towle Photos
Josiah H. Townsend Photos
Stanley Trauth Photos
Rick Treinen Photos
Dr. Amadej Trnkoczy Photos
Yi-jiun Tsai Photos
Michael W. Tuma Photos
B.S. Tuniyev Photos
Evan Twomey Photos
Kanishka Ukuwela Photos
Unknown (CAS) Photos
Jean-Pierre Vacher Photos
Javier Valera-Leal Photos
Ben Van Allen Photos
Jasper van Dalen Photos
Arie van der Meijden Photos
Tom Van Wagner Photos
Glenn and Martha Vargas Photos
Anand Varma Photos
Carlos R. Vasquez-Almazan Photos
Pierre-Yves Vaucher Photos
Jorge Vega Photos
Guillermo Velo Antón Photos
Miguel Vences Photos
Pablo Venegas Photos
Diogo Verissimo Photos
Museum of Vertebrate Zoology Photos
Martin Vestergaard Photos
David Vieites Photos
Luis J. Villanueva-Rivera Photos
Jens V. Vindum Photos
Rudolf von May Photos
Vance Vredenburg Photos
David Wake Photos
Marvalee H. Wake Photos
John W. Wall Photos
Henk Wallays Photos
Robert Wanat Photos
Tammy Ward Photos
Dee E. Warenycia Photos
Nick D. Waters Photos
Charles Webber Photos
Erik Wederkinch Photos
Edgar A. Wefer Photos
Sara Weinstein Photos
Dr. Peter Weish Photos
H. Welsh Photos
Frans Wennmacker Photos
Mette Westergaard Photos
John White Photos
Kellie Whittaker Photos
E. Wild Photos
Bert Willaert Photos
Christopher J. Williams Photos
Ryan Winkleman Photos
Doug Wirtz Photos
Germano Woehl Jr. Photos
David Wojnowski Photos
Sebastian Wolf Photos
Ron Wolf Photos
Wolfgang Wuster Photos
Victor Hugo Zaracho Photos
Yu Zeng Photos
Peng Zhang Photos
Haitao Zhao Photos
Jing Che and Haitao Zhao Photos
Doug Zimmerman Photos

Reviewing Distribution Data
Reviewed the distribution data for Indian species
César Barrio-Amorós Reviewed the distribution data for Venezuelan species
Reviewed the distribution data for Guatemalan species
Reviewed the distribution data for Hong Kong species
Reviewed the distribution data for Australian species
Nikolai Orlov Reviewed the distribution data for Vietnamese species
Reviewed the distribution data for Costa Rican species
Bryan Stuart Reviewed the distribution data for Cambodian and Lao species

Commercial institutions
Edition Chimaira publishers,
Frankfurt am Main
Kindly permitted the publication of material from "Treefrogs of Africa" by Arne Schiřtz (1999).

Academic institutions
and Dave Reddy
at the American Museum of Natural History
Provided and continue to provide up-to-date taxonomic data through the Amphibian species of the world database.
California Academy of Sciences
Help us answering incoming email, and provide a link to their specimen database.
Ellin Beltz at Northeastern Illinois University
Provided a link to the "Translations of the Scientific Names of the Reptiles and Amphibians of North America".
Field Museum of Natural History
Provide a link to their specimen database.
Museum of Comparative Zoology, Harvard University
Provide a link to their specimen database.
Peabody Museum of Natural History, Yale University
Provide a link to their specimen database.
Swedish Museum of Natural History
Provide a link to their specimen database.

Conservation organizations
IUCN, CI, and NatureServe
Provide links to, and text and map data from, the Global Amphibian Assessment.
NatureServe Explorer
Provide links to conservation statuses of U.S. and Canadian species.

Students who helped write species accounts during David Wake's 1998 Amphibian Population Declines Seminar
Lauren M. Chan Species accounts
Yair Chaver Species accounts
Erica Garcia Species accounts
Sarah Graber Species accounts
Amy Jess Species accounts
Alan Krakauer Species accounts
Rachel L. Mueller Species accounts
Sasha Norwood Species accounts

Students who helped write species accounts during James Hanken and David Wake's 2002 Amphibian Population Declines Seminar
Cailin O'Connor Species accounts
Charles P. McCormick Species accounts
Christopher Searcy Species accounts
Christopher Streeter Species accounts
D.C. Blackburn Species accounts
Mark D. Cooperman Species accounts
Michelle Knapp Species accounts
Phoebe Lehmann Species accounts
Ryan Kerney Species accounts

Contributions by students in the Harvard herpetology course, taught by Jim Hanken and Jonathan Losos, Fall 2008
John Abraham Species accounts
Amy Ru Chen Species accounts
Cindy Liu Species accounts
Kristina Prus Species accounts
Matan Shelomi Species accounts
Yunke Wu Species accounts

Contributions by students in the Black Hills State herpetology course, taught by Brian Smith, Fall 2008
Christine Bubac Species accounts
Nathaniel Diede Species accounts
Dustin Guericke Species accounts
Jodi Massie Species accounts
Jeni Maxon Species accounts
Mark Mazza Species accounts
Jason Nies Species accounts
Victoria Sprague Species accounts
Nathan Van Patten Species accounts

Contributions by the AmphibiaWeb Team
David Blackburn Family-level accounts, facilitating photo contribution from KU, systematics
David Cannatella Family-level accounts, systematics
Raul Diaz Species accounts
Rebecca Doubledee Species accounts
David Fisher Photo lists for North American amphibian species lacking images; georeferencing
Joyce Gross Photos, AmphibiaWeb and CalPhotos database development and maintenance
Michelle Koo Cartograms, georeferencing, new species graphs
Greg LaMonte Species accounts
Meredith J. Mahoney Species accounts
Arie van der Meijden Species accounts, Photos, website usage statistics
Hien Nguyen AmphibiaWeb Logo; Added maps & sound files
Brian Petirs Species accounts
April Robinson Species accounts
Franziska Sandmeier Species accounts
Sean Schoville Species accounts
Ambika Sopory Species accounts
Carol Spencer Inclusion of specimen localities on AmphibiaWeb's dynamic maps via HerpNET
Rachna Tiwari Editing Species accounts
Tate Tunstall Species accounts
Vance Vredenburg Species accounts, Photos
David Wake PI, systematics, new species>, species accounts, Photos
Kellie Whittaker Species accounts (new), Species accounts (edited), Photos
Nicole Wong Standardizing reference database

Contributions by Berkeley URAP students and summer interns
Catherine Aguilar Species accounts
Ketti Augusztiny Species accounts
Peera Chantasirivisal Species accounts
David Chen Species accounts
Anna Chow Species accounts
Nelly Chow Species accounts
Kenny D'Oyen Species accounts
Joseph Engeda New species georeferencing and literature searches
Benjamin Fryer Species accounts
Lettie Gallup Species accounts
Veronica Garza Species accounts
Anisha Gandhi Species accounts
Kevin Gin Species accounts
Kip Green Species accounts
Christine Isabel Ilagan Species accounts
Michelle Iwaki Species accounts
Sandya Iyer Species accounts
Taha Jabbar Species accounts
Zaina Khan Species accounts
Stella Yujin Kim Homepage redesign, Species accounts
Alexis Leigh Krup Species accounts
Deborah Lee Species accounts
Cheng (Lily) Li Species accounts
Michael Li Species accounts
Jesse Lou Species accounts
Vanessa Lovenburg Species accounts
Christine Lu Species accounts
Keith Lui Species accounts
Theresa Ly Species accounts
Raahil Malhotra Species accounts
Shelly Lyser Species accounts
Omar Malik Species accounts
Sijie Mao Species accounts
Janel Marcelino Species accounts
Prathna Mehta Species accounts
Asako Miyakawa Species accounts
Rupi Mudan Species accounts
Alamelu Natesan Species accounts
Gordon Ngo New species georeferencing and literature searches
Elizabeth Reisman Species accounts
Sunny Shah Species accounts
Greg Suryana New species georeferencing and literature searches
Gary Tsai Species accounts
Stephanie Ung Species accounts
Chih Wang Species accounts
Nichole Winters Species accounts
Arman Zaman Species accounts
Xi Zhai Species accounts
Henry Zhu Species accounts