Tarkhnishvili, D. N. and Gokhelashvili, R. K. (1999). ''The amphibians of the Caucasus.'' Advances in Amphibian Research in the Former Soviet Union, 4, 1-233.
Terent'ev, P. V. and Chernov, S. A (1965). Key to Amphibians and Reptiles [of the USSR]. Israel Program for Scientific Translations, Jerusalem.
Written by Sergius L. Kuzmin (ipe51 AT yahoo.com), Institute of Ecology and Evolution, Russian Academy of Sciences, Moscow
First submitted 2005-10-26
Edited by Sean Schoville (1/18/00), D. B. Wake (10/26/05) (2005-10-26)
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