Tarkhnishvili, D. N. and Gokhelashvili, R. K. (1999). ''The amphibians of the Caucasus.'' Advances in Amphibian Research in the Former Soviet Union, 4, 1-233.
Terent'ev, P. V. and Chernov, S. A (1965). Key to Amphibians and Reptiles [of the USSR]. Israel Program for Scientific Translations, Jerusalem.
Written by Sergius L. Kuzmin (ipe51 AT yahoo.com), Institute of Ecology and Evolution, Russian Academy of Sciences, Moscow
First submitted 1999-09-30
Edited by Vance Vredenburg, Kellie Whittaker (2008-09-19)
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